Authors

Dimitri Joseph

Type

Text

Type

Thesis

Advisor

Hollingsworth, Nancy M | Luk, Ed.

Date

2017-05-01

Keywords

Cellular biology | homologous recombination, srs2, zip1

Department

Department of Biochemistry and Cell Biology

Language

en_US

Source

This work is sponsored by the Stony Brook University Graduate School in compliance with the requirements for completion of degree.

Identifier

http://hdl.handle.net/11401/76890

Publisher

The Graduate School, Stony Brook University: Stony Brook, NY.

Format

application/pdf

Abstract

Homologous Recombination (HR) is an essential mechanism for double strand break (DSB) repair that generates crossovers (COs). In meiosis, HR is required for proper chromosome segregation at the first meiotic divison. The Sgs1 helicase promotes the formation of non-crossovers (NCOs) through a process called synthesis-dependent strand annealing. A set of meiosis specific genes called the ZMM genes, protects strand invasion intermediates from Sgs1, resulting in the formation of a specific class of COs that are distributed throughout the genome. The transverse filament protein, Zip1 is encoded by one of the ZMM genes, and phosphorylation of Zip1 on four adjacent serines in its C terminus are required for the ZMM pathway of recombination. Previous work has shown the absence of SGS1 combined with a nonphosphorylatable version of ZIP1, zip1-4A, triggers the meiotic recombination checkpoint, resulting in meiotic prophase arrest. Srs2 is a less well studied helicase in meiosis, which like Sgs1, exhibits an anti-recombinase function during DSB repair in vegetative cells. The goal of my thesis was to see whether depletion of SRS2 has a similar genetic interaction in zip1-4A, which would suggest a role for SRS2 during meiosis similar to SGS1. | 53 pages

Share

COinS
 
 

To view the content in your browser, please download Adobe Reader or, alternately,
you may Download the file to your hard drive.

NOTE: The latest versions of Adobe Reader do not support viewing PDF files within Firefox on Mac OS and if you are using a modern (Intel) Mac, there is no official plugin for viewing PDF files within the browser window.